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Except where otherwise noted, this item's license is described as Copyright the authors. Publisher PDF posted as allowed by the publisher's author right policy at https://www.molbiolcell.org/info-for-authors#license. Under the License and Publishing Agreement, authors grant to the general public, effective two months after publication of (i.e., the appearance of the edited manuscript in an online issue of MBoC), the nonexclusive right to copy, distribute, or display the manuscript subject to the terms of the Creative Commons Attribution 4.0 International (CC-BY 4.0) License (https://creativecommons.org/licenses/by/4.0).